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degeneration of muscles in hand and foot go on side by side, because some are used and some are disused. Centres of use and disuse must be centres of evolution. And there would be as many distinct centres of evolution in different parts of the body as there were centres of use and disuse. And between these centres there might be no correlation except that of use and disuse. Brain, muscles, and jaws would develop simultaneously in the ancestors of insects. And the effects of use and disuse, transmitted through a series of generations, would be cumulative. The species advances rapidly because all its members have in general the same habits; the same parts are advancing or degenerating, although at different rates, in all its individuals. An animal having an organ highly developed is far less likely to pair with one having a lower development of the same organ. The NeoLamarckian theory supplies thus what is lacking in the Neo-Darwinian.

In lower forms, like hydra, of simple structure and comparatively few possibilites of variation, natural selection is dominant. In higher forms, like vertebrates, and especially in man, it is of decidedly subordinate value as a promoter of evolution. For man, as we have seen, is a marvellously complex being. The great difficulty in his case is not so much to quickly gain new and favorable variations as to keep all the organs and powers of the body steadily advancing side by side. Natural selection has in man the important but subordinate position of the judge in a criminal court, to pronounce the death verdict on the hopeless and incorrigible.

Both Neo-Darwinians and Neo-Lamarckians have

erred in being too exclusively mechanical in their theories. It is the main business of the scientific man to discover and study mechanisms. But he must remember that mechanism does not produce force, it only transmits it. If he maintains that he has nothing to do with anything outside of mechanism, that the invisible and imponderable force lies outside of his domain, he has handed over to metaphysics the fairest and richest portion of his realm. In our fear of being metaphysical we have swung to another extreme, and have lost sight of valuable truth which lay at the bottom of the old vitalistic theories. Cells, tissues, and organs are but channels along which the flood of life-force flows. Boveri has well said, "There is too much intelligence (Verstand) in nature for any purely mechanical theory to be possible."

Each theory contains important truth. Nägeli's view of the importance of initial tendencies, inherent in the original living substance, is too often undervalued. My own conviction, at least, is steadily strengthening that, without some such original tendency or aim, evolution would never have reached its present culmination in man. His error lies in emphasizing this factor too exclusively. The fundamental proposition of Weismann's theory, that heredity is due to continuity of germ-plasm, seems to contain important truth. But we need not therefore accept his theory of a germ-plasm so isolated and independent as to be beyond control or influence by the habits of the body. The importance of use and disuse, and the transmissibility of their effects, would seem to supply a factor essential to evolution. Weismann has done good service in emphasizing the stability of the

germ-plasm. Evolution is always slow, and, for that very reason, sure.

If these conclusions are correct, they have an important practical bearing. Struggle and effort are essential to progress. Not inborn talent alone, but the use which one makes of it, counts in evolution. The effects of use and disuse are cumulative. The hard-fought battle of past generations becomes an easy victory in the present, just because of the strength acquired and handed down from the past struggle. Persistent variation toward evil is in time weeded out by natural selection. And, while evil remains in the world, we are to lay up stores of strength for ourselves and our descendants by sturdily fighting it. But the effects of right living through a hundred generations are not overcome by the criminal life of one or two. Evil surroundings weigh more in producing criminals than heredity, and their children are not irreclaimable.

The struggles and victories of each one of us encourage the rest. There is, to borrow Mr. Huxley's language, not only a survival of the fittest, but a fitting of as many as possible to survive. And in the midst of the hardest struggle there is the peace which comes from the assurance of a glorious triumph.

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Condensed Chart of Development of the Main Line of the Animal Kingdom leading to Man.

Annelid.

Primitive

Fish.

Amphibian.

Reptile.

Lower Placental Mammals.

Ape.

Man.

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